The year is 2258, ten years after the destruction of Earth by the Minbari fleet. The last refuge of Humanity is a devastated colony on the planet Proxima-3. President Mary Crane is the head of the Resistance Government. The remnants of the Earth Military continue to fight against the Minbari using raiding tactics. Relations with Narn allies are far from ideal. The last hope of Humanity is one single ship and one single person. This ship is Babylon, this man is its captain. John Sheridan.

The end of the beginning, or maybe the beginning of the end. Sheridan reaps the rewards of his earlier deeds, Delenn enters Chrysalis, Ivanova makes plans for the future, Sinoval searches for proof, Lord Refa visits Proxima 3, and Londo begins to realize that a serious threat looms over him.